All Publications

Peer-Reviewed Articles

Troupin D, Carmel Y. 2018. Conservation planning under uncertainty in urban development and vegetation dynamics. PLoS ONE 13(4): e0195429.

Shabtay A, Portman ME, Carmel Y. 2018. Contributions of marine infrastructures to marine planning and protected area networking.  Aquatic Conservation, 2018:1-10.

Shabtay, A., Portman, M. E., Carmel, Y. (2017), Incorporating principles of reconciliation ecology to achieve ecosystem-based marine spatial planning. Ecological Engineering. 10.1016/j.ecoleng.2017.08.026

Eshel G, Carmel Y. 2017. Expanded View of Ecosystem Stability: A Grazed Grassland Case StudyPLoSONE, 12(6) e0178235

Carmel, Y., Suprunenko, Y. F., Kunin, W. E., Kent, R., Belmaker, J., Bar-Massada, A. and Cornell, S. J. (2017), Using exclusion rate to unify niche and neutral perspectives on coexistence. Oikos. doi:10.1111/oik.04380

Makler-Pick, V., Hipsey, M., Zohary, T., Carmel, Y., and Gal, G. 2017. Intraguild predation dynamics in a lake ecosystem based on a coupled hydrodynamic-ecological model: the example of lake Kinneret (Israel). Biology 6(22), doi:10.3390/biology6020022.

Troupin D, Carmel Y. 2016. Landscape patterns of development under two alternative scenarios: implications for conservation.   Land Use Policy 54:221-234.

Nezer O, Bar-David S, Gueta, T, and Carmel, Y. 2016. High-resolution species-distribution model based on systematic sampling and indirect observations. Biodiversity and Conservation. doi:10.1007/s10531-016-1251-2

Gueta T. and Carmel, Y. 2016. Quantifying the value of user-level data cleaning for big data: A case study using mammal distribution modelsEcological Informatics 34: 139–145.

Liberzon J, Carmel Y and Angel DL. 2016. The elusive nature of fluorescent organic matter in Lake Kinneret, Israel. Hydrobiologia 763 (1), 53-68.

Kent R, Bar-Massada A, and Carmel, Y. 2014. Bird and mammal species composition in distinct geographic regions and their relationships with environmental factors across multiple spatial scales. Ecology and Evolution, 4(10):1963-1971.

Bar-Massada A, Kent R, and Carmel, Y. 2014. Environmental heterogeneity affects the location of modeled communities along the niche-neutrality continuum. Proceedings of the Royal Society B, 281:20133249.

Troupin, D. and Carmel, Y. 2014. Can agro-ecosystems efficiently complement protected area networks? Biological Conservation, 169:158-166.

Blank, L., Linker, R. and Carmel, Y. 2013. A Multiscale Analysis of Herbaceous Species Richness in a Mediterranean Ecosystem. Journal of Plant Ecology, 6(2):113-121.

Sutherland, W.J., Freckleton, R.P., Godfray, H.C.J., Beissinger, S.R., Benton, T., Cameron, D.D., Carmel, Y., Coomes, D.A., Coulson, T., Emmerson, M.C., Hails, R.S., Hays, G.C., Hodgson, D.J., Hutchings, M.J., Johnson, D., Jones, J.P.G., Keeling, M.J., Kokko, H., Kunin, W.E., Lambin, X., Lewis, O.T., Malhi, Y., Mieszkowska, N., Milner-Gulland, E.J., Norris, K., Phillimore, A.B., Purves, D.W., Reid, J.M., Reuman, D.C., Thompson, K., Travis, J.M.J., Turnbull, L.A., Wardle, D.A. and Wiegand, T. 2013. Identification of 100 fundamental ecological questions. Journal of Ecology, 101(1):58-67.

Carmel, Y., Kent, R., Bar Massada, A., Blank, L., Liberzon, J., Nezer, O., Sapir, G. and Federman, R. 2013. Trends in ecological research during the last three decades – a systematic review. PlosONE, 8(4):e59813.

Davidson, A., Carmel, Y. and Bar David, S. 2013. Characterizing movement pathways using a non-invasive approach: The reintroduced Asiatic wild ass in the Negev. Landscape ecology, 28(8):1465-1478.

Federman, R., Kent, R. and Carmel, Y. 2013. Irrigation as an important factor in species distribution models. Basic and Applied Ecology, 14(2013) 651–658.

Bar Massada, A., Kent, R., Blank, L., Perevolotsky, A., Hadar, L. and Carmel, Y. 2012. Automated segmentation of vegetation structure units in a Mediterranean landscape. International Journal of Remote Sensing, 33(2):346-364.

Blank, L. and Carmel, Y. 2012. Woody vegetation patch types affect herbaceous species richness and composition in a Mediterranean ecosystem. Community Ecology, 13(1):72-81.

Kent, R., Bar Massada, A. and Carmel, Y. 2011. Multiscale analyses of mammal species composition – environment relationship in the contiguous USA. PLoSONE, 6(9):e25440.

Gabay, O., Perevolotsky, A., Bar Massada, A., Carmel. Y. and Shachak, M. 2011. Differential effects of goat browsing on herbaceous plant community in a two-phase mosaic. Plant Ecology, 212(10):1643-1653.

Paz, S., Carmel, Y., Jahshan, F. and Shoshany, M. 2011. Post-fire Analysis of Pre-fire mapping of fire-risk: A recent case study from Mt. Carmel (Israel). Forest Ecology and Management, 262(7):1184-1188.

Kent, R. and Carmel, Y. 2011. Presence-only vs. Presence-absence data in species composition determinant analyses. Diversity & Distributions, 17:474-479.

Makler, V., Gal, G., Gorfine, M., Hipsey. M, and Carmel, Y. 2011. Sensitivity analysis for complex ecological models – a new approach. Environmental Modelling & Software, 26:124-134.

Kent, R. and Carmel, Y. 2011. Evaluation of five clustering algorithms for biodiversity surrogates. Ecological Indicators, 11:896-901.

Eitan, O., Barchana, M., Linn, S., Yuval., Dubnov, J., Carmel, Y. and Broday D. 2010. Spatial analysis of air pollution and cancer incidence rates in Haifa Bay, Israel. Science of the Total Environment, 408(20):4429-4439.

Van Leeuwen, W.J.D., Casady, G.M., Neary, DG., Bautista, S., Alloza, J.A., Carmel, Y., Wittenberg, L., Malkinson, D. and Orr, B.J. 2010. Monitoring post-wildfire vegetation response with remotely sensed time-series data in Spain, USA and Israel. International Journal of Wildland Fire, 19:75-93.

Bar Massada, A., Carmel, Y., Koniak, G. and Noy Meir, I. 2009. The effects of disturbance based management on the dynamics of Mediterranean vegetation: A hierarchical and spatially explicit modeling approach. Ecological Modelling, 220:2525-2553.

Dolev, A. and Carmel, Y. 2009. Distribution of threatened–unprotected vertebrates as a basis for conservation planning. Israel Journal of Ecology and Evolution, 55(2):117-132.

Steele, K., Carmel, Y., Cross, J. and Wilcox, C. 2009. Uses and Misuses of Multi-Criteria Decision Analysis (MCDA) in Environmental Decision-Making. Risk Analysis, 29(1):26-33.

Carmel, Y., Paz, S., Jahshan, F. and Shoshany, M. 2009. Assessing fire risk using Monte Carlo simulations of fire spread. Forest Ecology and Management, 257:370-377.

Bar Massada, A., and Carmel, Y. 2008. Incorporating output variance in local sensitivity analysis for stochastic ecological models. Ecological Modelling, 213:463-467.

Bar Massada, A., Gabay, O., Perevolotsky, A., and Carmel, Y. 2008. Quantifying the effect of grazing and shrub clearing on small scale spatial pattern of vegetation. Landscape Ecology, 23:327-339.

Grünzweig, J.M., Carmel, Y., Riov, J., Sever, N., McCreary, D.D., Flather, C.H. 2008. Growth, resource storage, and adaptation to draught in California and eastern Mediterranean oak seedlings. Canadian Journal of Forest Research, 38:331-342.

Oman, S.D., Landsman, V., Carmel, Y. and Kadmon, R. 2007. Analyzing spatially distributed binary data .using independent-block estimating equations. Biometrics, 63:892-900.

Segal, B., Sapir, Y. and Carmel, Y. 2006. Fragmentation and pollination crisis in the self incompatible Iris bismarckiana (iridaceae), with implications for conservation. Israel Journal of Ecology & Evolution, 52:111-112.

Carmel, Y. and Stoller-Cavari, L. 2006. Comparing environmental and biological surrogates for biodiversity at a local scale. Israel Journal of Ecology & Evolution, 52:11-27.

Bar Massada, A., Carmel, Y., Even Tzur, G., Grünzweig, J.M. and Yakir, D. 2006. Assessment of temporal changes in aboveground forest tree biomass using aerial photographs and allometric equations. Canadian Journal of Forest Research, 36(10): 2585-2594.

Moilanen, A., Runge, M.C., Elith, J., Tyre, A., Carmel, Y., Fegraus, E., Wintle, B., Burgman, M. and Ben-Haim, Y. 2006. Planning for robust reserve networks using uncertainty analysis. Ecological Modelling, 199:115-124.

Carmel. Y. and Flather, C. H. 2006. Constrained Range Expansion and climate change assessments. Frontiers in Ecology and the Environment. 4(4):178-179. Peer-Reviewed Letter.

Carmel, Y. and Ben-Haim, Y. 2005. Info-Gap Robust-Satisficing Model of Foraging Behavior: Do Foragers Optimize or Satisfice? The American Naturalist. 166 (5):633-641.

Yuval., Broday, M. D. and Carmel, Y. 2005. Mapping Spatio-Temporal Variables: The Impact of the Time-Averaging Window Width on the Spatial Accuracy. Atmospheric Environment, 39:3611-3619.

Svoray, T. and Carmel, Y. 2005. Empirical Method for Topographic Correction in Aerial Photographs. IEEE Geoscience and Remote Sensing Letters, 2 (2):211-214.

Altman, T., Carmel Y., Gueta, R., Zaslavsky D. and Doytsher Y. 2005. Assessment of an “Energy Tower” Potentials in Australia Using a Mathematical Model and GIS. Solar Energy, 78:799-808.

Carmel, Y. and Flather, C.H. 2004. Comparing Landscape Scale Vegetation Dynamics Following Recent Disturbance in Climatically Similar Sites in California and the Mediterranean Basin. Landscape Ecology. 19: 573-590.

Carmel, Y. 2004. Controlling Data Uncertainty via Aggregation in Remotely Sensed Data. IEEE Geoscience and Remote Sensing Letters, 1 (2):39-41.

Carmel, Y. 2004. Characterizing Location and Classification Error Patterns in Time-Series Thematic Maps. IEEE Geoscience and Remote Sensing letters, 1 (1):11-14.

Carmel, Y. and Dean, D.J. 2004. Performance of a Spatio-Temporal Error Model for Raster Datasets under Complex Error Patterns. International Journal of Remote Sensing. 25 (23): 5283-5296.

Carmel, Y. and Naveh, Z. 2002. The Paradigm of Ecosystem and the Paradigm of Landscape in Ecology – Implications for Land Planning and Management in the Mediterranean Region. Journal of Mediterranean Ecology, 3(2-3):35-46.

Carmel, Y., Dean, D.J. and Flather, C.H. 2001. Combining Location and Classification Error Sources for Estimating Multi-Temporal Database Accuracy. Photogrammetric Engineering & Remote Sensing, 67 (7): 865-872.

Carmel, Y., Kadmon R. and Nirel, R. 2001. Spatio-Temporal Predictive Models of Mediterranean Vegetation Dynamics. Ecological Applications, 11 (1):268-280.

Carmel, Y. and Kadmon, R. 1999. Effects of Grazing and Topography on Long Term Vegetation Changes in a Mediterranean Ecosystem in Israel. Plant Ecology, 145:239-250.

Carmel, Y., Kadmon, R. 1998. Computerized Classification of Mediterranean Vegetation Using Panchromatic Aerial Photographs. Journal of Vegetation Science, 9:445-454.

Carmel, Y. and Safriel, U.N. 1998. Habitat Use by Bats in a Mediterranean Ecosystem in Israel – Conservation implications. Biological Conservation, 84:245-250.


Agra, H., Carmel, Y., Smith, R.K. and Ne’eman, G. (2016) Forest conservation: Global evidence for the effects of interventions. University of Cambridge, Cambridge, UK

Book Chapters

Agra, H., Carmel, Y., Smith, R.K. and Ne’eman, G. (2016) Forest conservation: Global evidence for the effects of interventions. University of Cambridge, Cambridge, UK

Carmel, Y. 2005. Aggregation as a Means of Increasing Thematic Map Accuracy. In: Atkinson, P, Martin, D, and Foody, GM, editors. Geodynamics. CRC Press, London. 29-38.

Naveh, Z., Carmel, Y. 2004. The Evolution of the Cultural Mediterranean Landscape in Israel as Affected by Fire, Grazing, and Human Activities. In: Wasser, S., editor. Evolutionary theory and processes: modern horizons. Kluwer Academic Publishers, Dordrecht, The Netherlands. 337-409.

Carmel, Y., Kadmon, R. 2000. Using Historical Aerial Photographs for Mapping Vegetation Changes. In: Kedar, B., editor. Remote Sensing. Yad Ben Zvi, Jerusalem, Israel. 77-83. In Hebrew.

Muskin (Carmel), Y. 1994. Habitat Use and Foraging Behavior of Insectivorous Bats in the Mediterranean Region. In: Shalmon, B., Simon, D. and Barak, Y. Editors. Bats of Israel. The Society for Protection of Nature in Israel. 110-119. In Hebrew.

Muskin (Carmel), Y. 1994. Bats in Mt. Meron. In: Markus, M., ed. Mount Meron – a landscape survey. Nature Reserve Authority, Jerusalem. 165-168. In Hebrew.

Book Reviews

Carmel, Y. 1997. Recent Advances in the Study of Bats. Global Ecology and Biogeography letters 6 (6): 453-454.


Lortet iris conservation: establishment of new populations of the endemic Lortet iris (Iris lortetii) (a progress report). 2014. In Hebrew

Sapir,N.and Carmel, Y. 2010. Post fire vegetation regeneration in planted pine forests. Ecology & Environment. 3:14-23. In Hebrew.

Mount Carmel: Risk Map vs. 2010 Fire (a map)

Nezer,O.and Carmel, Y. 2010. Automated mapping of Acacia trees in the Arava valley using remote sensing (a report). In Hebrew.

Judaean Mountains fire risk report (2010). In Hebrew

Mount Carmel area fire risk report (2004-2007). In Hebrew


Bar Massada,A. (2008), Mapping and modeling the dynamics of Mediterranean vegetation under various management activities, (PhD dissertation)

Vdov,O. (2013), The linkage between environmental heterogeneity and biodiversity in algal assemblages on the rocky shore, (Master dissertation)

Kent,R. (2010), Relationships between environmental parameters and species composition patterns and their implications for biodiversity surrogates – a multiple scale study, (PhD dissertation)

Nezer,O. (2011), Modeling the distribution of the Asiatic Wild Ass (Equus Hemionus) in the Negev Desert, Southern Israel, (Master dissertation)

Blank,L. (2009), Multiscale analysis of species distriburion pattern(PhD dissertation)

Federman,R. (2010), Potential distribution and monitoring the invasive species “The little fire ant” (Wassmania auropunctata), (Master dissertation)

Liberzon,J. (2011) Seasonal dynamics of chromophoric dissolved organic matter in Lake Kinneret: An investigation of fluorescent component changes in relation to microbial and phytoplankton productivity, (Master dissertation)

Segal,B. (2006), Nature Reserve Management for Conservation of the Endangered Species Iris bismarckiana, (Master dissertation)

Jahshan,F. (2010), Mapping fire risk using multiple simulations of fire behavior, (Master dissertation)

Noi Harpaz,G. (2014), Development of a quantitative approach for Environmental Impact Assessment of a desalination plan: Shavei-Zion case study, (Master dissertation)


Carmel, Y., Flather, C., Dean, D. 2006. A Methodology for Translating Positional Error into Measures of Attribute Error, and Combining the Two Error Sources. A Plenary Lecture in: Proceedings of the Seventh International Symposium on Spatial Accuracy Assessment in Natural Resources and Environmental Sciences, Lisbon.